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学习笔记 | 长期记忆的生物学基础

  • 2026-02-10 08:26:54
学习笔记 | 长期记忆的生物学基础

学习笔记 | 长期记忆的生物学基础

​核心定义​：长期记忆指生物在睡眠后仍能保留的记忆，是神经系统动物的普遍特征，但不同物种记忆能力存在显著差异（如线虫、金鱼常被误认为无长期记忆）。

​物质基础​：长期记忆的形成依赖于8种关键蛋白质及其对应基因​的协同作用，它们通过调节神经突触的可塑性实现记忆存储。

🧠 8种记忆蛋白功能详解

1. NMDAR蛋白质
功能：神经刺激感应器
-对各种神经刺激进行筛选，识别值得记忆的重要事件
-充当"大门守卫"，只让重要信号通过
-控制钙离子进入神经元，启动记忆形成过程

特点：非特异性，参与多种神经活动。

2. CaMKII蛋白质
功能：神经元兴奋维持器
-维持神经元的持续兴奋状态
-促进神经元向外释放信号
-确保记忆信号能够持续传递

特点：非特异性，影响神经信号传递

3. CREB蛋白质
功能：信号传递与批准者
-接收来自CaMKII的信号
-激活并批准释放Arc蛋白
-作为连接上游信号和下游执行的关键桥梁

4. Arc蛋白质
功能：长期记忆核心执行者
-呈空心球状结构，携带mRNA遗传信息
-是唯一专用于长期记忆的蛋白质
-在突触重构中发挥核心作用

特点：具有病毒感染特性，是生物演化的"奢侈品"

5. mTOR蛋白质
功能：突触构建启动器
-与Arc蛋白相遇后启动突触构建工作
-召唤Actin蛋白参与施工
-调控神经通路的加宽和加强

6. Actin蛋白质
功能：神经通路施工者
-在mTOR调控下加宽神经通路
-实际执行突触连接的增强工作
-让神经通路变得更加粗壮稳固

7. PKMζ蛋白质
功能：突触保护者
-负责保护新建的突触结构
-维护记忆的稳定性
-与Rac1蛋白形成竞争关系

8. Rac1蛋白质
功能：突触修剪者
-负责修剪和弱化神经突触
-与PKMζ形成"记忆竞争"关系
-胜出时导致记忆遗忘，PKMζ胜出时记忆保留

协同工作机制总结
这8种蛋白形成了完整的记忆形成链条：
筛选感应 → 信号维持 → 信号传递
核心执行 → 构建启动 → 实际施工
保护维护 ↔ 修剪竞争

唯一特殊蛋白：Arc蛋白是唯一仅参与长期记忆的"奢侈品"，其他7种蛋白在基础生命活动中也必不可少。

这个系统展现了生物学上精妙的分工合作，其中Arc基因被称为生物演化的"奢侈品"和"生命游戏DLC"，其起源可能是远古病毒感染真核生物后遗留的基因片段。这一"病毒遗产"赋予高级生物高效长期记忆能力，成为人类文明形成的关键神经基础。#神经科学入门

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  128. /yingpanguazai/ssd/ssd1/www/no.67808.cn/vendor/topthink/think-helper/src/Collection.php ( 16.47 KB )
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  134. /yingpanguazai/ssd/ssd1/www/no.67808.cn/vendor/topthink/think-view/src/Think.php ( 8.38 KB )
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  1. CONNECT:[ UseTime:0.000457s ] mysql:host=127.0.0.1;port=3306;dbname=no_67808;charset=utf8mb4
  2. SHOW FULL COLUMNS FROM `fenlei` [ RunTime:0.000535s ]
  3. SELECT * FROM `fenlei` WHERE `fid` = 0 [ RunTime:0.000256s ]
  4. SELECT * FROM `fenlei` WHERE `fid` = 63 [ RunTime:0.000264s ]
  5. SHOW FULL COLUMNS FROM `set` [ RunTime:0.000471s ]
  6. SELECT * FROM `set` [ RunTime:0.000188s ]
  7. SHOW FULL COLUMNS FROM `article` [ RunTime:0.000589s ]
  8. SELECT * FROM `article` WHERE `id` = 462129 LIMIT 1 [ RunTime:0.001940s ]
  9. UPDATE `article` SET `lasttime` = 1770842076 WHERE `id` = 462129 [ RunTime:0.005696s ]
  10. SELECT * FROM `fenlei` WHERE `id` = 65 LIMIT 1 [ RunTime:0.000303s ]
  11. SELECT * FROM `article` WHERE `id` < 462129 ORDER BY `id` DESC LIMIT 1 [ RunTime:0.001140s ]
  12. SELECT * FROM `article` WHERE `id` > 462129 ORDER BY `id` ASC LIMIT 1 [ RunTime:0.000398s ]
  13. SELECT * FROM `article` WHERE `id` < 462129 ORDER BY `id` DESC LIMIT 10 [ RunTime:0.007614s ]
  14. SELECT * FROM `article` WHERE `id` < 462129 ORDER BY `id` DESC LIMIT 10,10 [ RunTime:0.011565s ]
  15. SELECT * FROM `article` WHERE `id` < 462129 ORDER BY `id` DESC LIMIT 20,10 [ RunTime:0.009934s ]
0.111303s